Nicholas A. Friedenberg, Ph.D

Vice President and

Research Director for Agriculture and Ecotoxicology

Nicholas A. Friedenberg joined Applied Biomathematics in 2007 and has since led projects on a wide range of subjects including endangered and invasive species, wind energy, bio-terrorism, forest pests, and pesticide resistance. His research and development activities have been supported by grants from USDA, DHS, SERDP, and Monsanto, as well as contracts from EPRI, and USACE. Nick serves clients in both the public and private sector.

Nick currently leads the company's activities in ecotoxicology and agricultural applications. He is one of the authors, along with Kevin Shoemaker, of the RAMAS IRM software package for modeling the evolution of insect pest resistance to genetically modified crops.

Nick received his B.A. in biology from Carleton College and his Ph.D. from Dartmouth College's Department of Ecology and Evolutionary Biology. His publications include both evolutionary and ecological studies.


Selected publications
More complete list maintained here.

Friedenberg, N.A.. 2020. Population-level risks to hoary bats amid continued wind energy development: assessing fatality reduction targets under broad uncertainty. Electric Power Research Institute, Palo Alto, CA. EPRI website

Kinlock, N.L., A.J. Laybourn, C.E. Murphy, J.J. Hoover, N.A. Friedenberg. 2020. Modelling bioenergetic and population-level impacts of invasive bigheaded carps (Hypophthalmichthys spp.) on native paddlefish (Polyodon spathula) in backwaters of the lower Mississippi River. Journal of Freshwater Biology 00:1-15. Journal website

Friedenberg, N.A., and C.M. Foley. 2018. From wind to wires: insight into a developing framework for incidental take permits for eages. Electric Power Research Institute, Palo Alto, CA. EPRI website

Trump, B.D., C. Foran, T. Rycroft, M.D. Wood, N. Bandolin, M. Cains, T. Cary, F. Crocker, N.A. Friedenberg, P. Gurian, K. Hamilton, J.J. Hoover, C. Meyer, K. Pokrzywinski, R. Ritterson, P. Schulte, C. Warner, E. Perkins, I. Linkov. 2018. Development of community of practice to support quantitative risk assessment for synthetic biology products: contaminant bioremediation and invasive carp control as cases. Environment Systems and Decisions. pdf

Martinez, J.C., M.A. Caprio, and N.A. Friedenberg. 2018. Density dependence and growth rate: evolutionary effects on resistance development to Bt (Bacillus thuringiensis). Journal of Economic Entomology 111: 382-390. pdf

Monzón, J.D., and N.A. Friedenberg. 2018. Metrics of population status for long-lived territorial birds: a case study of golden eagle demography. Biological Conservation 220: 280-289. pdf

Friedenberg, N.A., J.J. Hoover, K. Boysen, and K.J. Killgore. 2018. Estimating abundance without recaptures of marked pallid sturgeon in the Mississippi River. Conservation Biology 32:457-465 pdf

Powell, J.A., M.J. Garlick, B.J. Bentz, N.A. Friedenberg. 2018. Differential dispersal and the Allee effect create power-law behavior: distribution of spot infestations during mountain pine beetle outbreaks. Journal of Animal Ecology 87:73-86. pdf

Friedenberg, N., and C. Foley. 2016. Defining protection of endangered species from an integrated multispecies perspective. Technical Report 3002008414. Electric Power Research Institute, Palo Alto, CA. report

Friedenberg, N.A., and K.T. Shoemaker. 2014. RAMAS IRM version 2.1: software for risk-based durability assessment. Applied Biomathematics, Setauket, NY. more info

Thomas, M.J., M.L. Peterson, N. Friedenberg, J.P. Van Eenennaam, J.R. Johnson, J.J. Hoover, A.P. Klimley. 2013. Stranding of spawning run green sturgeon in the Sacramento River: post-rescue movements and potential population-level effects. North American Journal of Fisheries Management 33:287-297 pdf

Nicholas Friedenberg on Google Scholar

Nick's ResearchGate profile

Nick Friedenberg's publications at


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